STATE COLLEGE, Pa. -- Penn State went 80 yards in 23 seconds to send its game against No. 18 Michigan into overtime Saturday.
Then the real game started.
The Nittany Lions and Wolverines went back and forth for four overtimes, squandering as many opportunities as they cashed in on, before Bill Belton's 2-yard touchdown run gave Penn State a 43-40 win in Beaver Stadium.
Michigan kicker Brendan Gibbons made two field goals and missed two on the Wolverines' four possessions, and Penn State had one Sam Ficken make, one miss and a lost fumble before Belton's score.
Penn State (4-2, 1-1 Big Ten) won for the second time in overtime under Bill O'Brien, and Michigan (5-1, 1-1) lost for the first time this season.
Penn State trailed in the final minute of regulation before quarterback Christian Hackenberg hit Allen Robinson for a 14-yard gain that was initially ruled out-of-bounds but then overturned by replay, then found Brandon Felder for 29 yards, then again went to Robinson, who came down with a jump ball at the half-yard line. Hackenberg sneaked the ball in on the next play to tie the score, and Gibbons' 52-yard field goal attempt fell short.
Penn State's defense, which had forced six turnovers in its first five games, came up with three takeaways in the first half alone, and the Nittany Lions capitalized on two of them. Hackenberg threw a pass off his back foot to the back of the end zone to Felder following an interception of Devin Gardner by Jordan Lucas. Hackenberg threw a 20-yard touchdown to tight end Jesse James after defensive end Anthony Zettel had dropped into coverage and intercepted Gardner, who completed only 6 of 12 first-half passes, early in the second quarter.
The Nittany Lions added their final score of the half when Hackenberg scrambled on third down and rifled a 24-yard touchdown pass to Felder, who had missed the previous week's game against Indiana with an ankle injury.
Both teams struggled to run the ball in the first half. Michigan had 23 carries for 56 yards and Penn State had 17 carries for 38 yards.
Michigan, which had not beaten Penn State since 2007, got a score in the opening 10 seconds of the second half when Frank Clark recovered a Zach Zwinak fumble and returned it for a touchdown.
Two series later, Hackenberg was intercepted by Raymon Taylor, leading to a short Gibbons field goal that cut the lead to 21-20 with 8:50 left in the third quarter.
NOTES: Michigan offensive tackle Taylor Lewan left the game in the second quarter with an unspecified injury and did not return in the second half. ... It was officially the longest game in Penn State history. ... Penn State's victory was its fourth straight over Michigan, which had won each of the previous nine meetings.